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Unless your destination device’s write speed is higher than your source device’s read speed, then the difference isn’t going to matter. But when you format a partition, you are given a choice as to allocation unit size. Larger sizes are generally better optimized for larger files because more data can be read and written with fewer I/O requests, which is why they’re sometimes used for volumes that will host databases. The tradeoff is that they can be less efficient, because the minimum footprint of any given file on the volume will be a single allocation unit, and you also can’t have multiple files occupying a single allocation unit. So if you have a 64 KB allocation unit (for example), then a 4 KB file will consume 64 KB, and a 65 KB file will consume 128 KB. The result is that you end up with more space lost to “slack” compared to smaller allocation unit sizes.
All that said, again I don’t know if tuning the allocation unit size will make much difference even if you don’t have the write speed constraint I described above. And I haven’t seen an official Macrium recommendation. Until recently, Image Guardian could only protect backups on NTFS volumes, but it now supports exFAT too.